Tuesday & Thursday, 8:30-9:50 am

Instructor: Dr. M.J. Crawford, Rm 215 Biological Sciences

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03 55 355 Embryology

Course Description:
        An amazing diversity of strategies are employed throughout the plant and animal kingdoms to build, to organize and to grow tissues, organs, and bodies.  Despite the various environmental niches inhabited, and the various morphologies that result, recent work has revealed that many of the genetic and mechanical means employed to achieve growth and change are shared in animals as diverse as flies and human.  By the end of this course you will come to understand how a body plan is established; how the mechanisms that give rise form are used by very different organisms to achieve similar ends; and how genetic networks have evolved a modularity that permits them to be employed over and over again throughout both evolution and development.  We will try to investigate these phenomena by exploring them through the experimental, observational, surgical, and genetics tools employed by scientists in the field.

Lecture meetings:     Section 01 8:30-9:50, Tuesday, Thursday  Odette 112

                                    Section 02 1:00-2:20 Tuesday, Thursday Dillon Hall Room: 264

Course Text

Principles of Development

Fifth Edition
Lewis Wolpert, Cheryll Tickle, and Alfonso Martinez Arias
Publication Date - May 2015
ISBN: 9780198709886
652 pages

Course Reserves

Wolpert et al., (2015) Principles of Development, 5th edition. (TBA)

Gilbert, Scott F. (2014) Developmental Biology, 10th ed.  Available at Leddy Library Course Reserves - 2 Hour Loan - Main Bldng - Circulation Desk ZPR030 .B001

Gilbert, Scott F. (2014) Developmental Biology, 9th ed.  Available at Leddy Library Course Reserves - 2 Hour Loan - Main Bldng - Circulation Desk QL 955 .G48 2010

Required Tools

Coloured pens or pencils

Instructors and Contacts

Graduate Teaching Assistants

Mohamad Fakhereddin,  Rm 111 Biological Sciences (morning section); Ingrid Qemo: (afternoon section)


Dr. Michael J. Crawford, Rm 111 Biological Sciences

Office Hours: 10:00 to11:00 Tuesday and Thursday, or by appointment.

Grading Scheme

Written Assignments                                   Due  Dates                                   

5%  bibliography                                  Sept 20, 2016                                  

Draft of DevoDictionary Page              Sept 29, 2016                                  

15% Editorial Comments                      Oct 6, 2016                                  

25% DevoDictionary Page                    Nov 19, 2016        

Mid-term Exam 25%                             Oct. 20, 2016        

Final Exam  30%   (tentative date)         Dec. 19, 2016 7:00 pm (tentative)

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