Thinking is the hardest work in
the world;
and most of us will go to great lengths to avoid it
Louise Dudley
Who am I? I was born in England
but worked mainly in Switzerland. I received the 1984 Nobel prize
for my theoretical work on the immune system. I developed the
network theory of antibody production and described the selective
theory of antibody formation
- Cytotoxic
T Lymphocytes
- Cytotoxic T lymphocytes "in action" as shown
at the Cells Alive! site.
URL -->
http://www.cellsalive.com/ctl.htm
- Plasma cell differentiation
- Information and images about plasma cell
differentiation at a site in the Department of
Parasitology (Immunology), Faculty of Medicine, Yamaguchi
University, Ube City, Japan.
URL -->
http://www.sv.cc.yamaguchi-u.ac.jp/~plasma/msg/nextpage/plasma04.html
Cell mediated and Humoral
Effector Responses: on line graphics presentation
CHAPTER 16: pp 379-412
- Effector Responses of Cell Mediated Immunity
- Properties of effector T cells
- activation requirements
- cell adhesion molecules
- effector molecules
- Direct cytotoxic responses
- CTL mediated cytotoxicity
- NK cell cytotoxicity
- antibody dependent cell mediated cytotoxicity
- experimental methods
- Delayed type hypersensitivity(DTH)
- phases of DTH response
- cytokines in DTH
- protective role
- detection of DTH
- pathological responses of DTH
- Effect Responses of Humoral Immunity
- Primary & secondary responses
- Experimental methods
- hemolytic plaque assay
- Elispot assay
- Hapten-carrier conjugates
- Regulation of Immune Effector Response
- antigen mediated regulation
- antibody mediated suppression
- immune complexes as regulators
- idiotypic regulation
- neuroendocrine regulation
DEFINE THE FOLLOWING KEY TERMS
clonal anergy |
clonal deletion |
immunological tolerance |
tolerogen |
Suppressor T cells |
|
ON COMPLETION OF THIS SECTION THE STUDENT SHOULD BE ABLE TO:
- Define and conceptualize immunological tolerance
- Describe and experiment that shows tolerance indcution
- Outline the mechanisms that induce self tolerance
- Briefly discuss the three phases of an immune response.
- Which phase is most complex and why?
- In which phase does antigen have little importance and
why?
- What distinguishes humoral immunity from cell mediated
immunity?
- What do we mean by a primary and a secondary immune
response?
- What characteristics usually distinguish the two
responses?
- Briefly discuss the antigen elimination curve.
- What things affect the shape of the curve and how does
this affect antibody production?
- Briefly describe the four phases of a primary antibody
response.
- When comparing the primary and secondary antibody
responses, three things change during the secondary
antibody response.
- What are they?
- Which one do you consider the most important? Why?
- Early experiments showing the importance of lymphocytes
to immunity followed two approaches.
- What were they?
- How did they differ?
- What were some of the initial observations that arose
from these two approaches?
- Describe the carrier effect.
- How did the study of the carrier effect resolve whether T
and B cells needed to react specifically with the same
epitope on an antigen molecule, and whether T cells and B
cells are specific for distinct antigenic determinants on
a complex antigen?
- Claman and his collegues provided the first direct
evidence that, for antibody responses, synergy was
required between T cells and B cells.
- Describe the experiment.
- Did the experiment show which cell did what? Explain. see
the following for details
- Claman, H. N., E.A. Chaperon and R.F. Triplett (1966)
Thymus marrow cell combinations. Synergism in antibody
production. Proc. Soc. Exp. Biol Med. 122:1167-1171
- Two kinds of immune cells are needed for antibody
responses.
- Mitchell and Miller's experiment demonstrated which cell
did what. How could they do this experiment if they did
not have antibodies specific for the two immune cells?
- Describe how Mitchison used the idea of the carrier
effect to determine whether the interacting T and B cells
recognized the same or different epitopes on a complex
antigen.
- Design an experiment proving that B cells recognize the
haptenic portion of a hapten carrier complex and that T
cells provide help in this interaction.
- B cells can internalize and process antigen.
- This fact resolves the following question:
"How could T helper cells with one receptor for both
an antigen fragment and an MHC-II molecule on the surface
of a macrophage provide antigen-specific help to a B cell
bearing antigen in its antibody receptor?"
- How do macrophages contribute to antibody production?
- How can the mixed lymphocyte reaction and the
cytotoxicity reaction be used to explain how T cells help
other T cells and which cell provides help and which cell
mediates cytotoxicity?
- Helper T cells can be divided into two subsets based on
their lymphokine secretion profile. Explain.
- As an immune response progesses, it needs to limit itself
and decrease in intensity. How does antibody feedback
accomplish this?
- Niels Jerne states that "antibodies recognize"
and are "being recognized" explain this
statement in terms of the network hypothesis.
- Briefly describe the dowregulatory cross regulation
between certain cytokines during the humoral immune
response.
- How does immunodificiency differ from immunotolerance?
- Design an experiment that shows tha "In utero"
or "in ovo" exposure to forein antigens leads
to tolerance in the adult animal.
- The clonal selection theory lends support to the idea of
immunologic tolerance. Explain.
- Briefly discuss some of the conditions that affect the
initiation of immunologic tolerance.
- What is common among these conditions?
- What are some of the basic differences between an antigen
and a tolerogen?
- Tolerance of self- antigens seems to be permanent, while
experimentally induced tolerance is not. Explain.
- The idea of clonal deletion suggests that self reactive
cells are eliminated (no cells to react to self antigens;
therefore, we tolerant self antigens), yet autoimmune
diseases occur (react to self antigens). Explain.